Migration of the ITCZ
Annual-mean precipitation distributions of four representative periods are shown in
Fig. 1 (continent configurations and precipitation distributions of all periods are in fig. S2 and movie S1). At 540 Ma, the supercontinent Pannotia had broken into Gondwana and several smaller continents (
Fig. 1A). From 540 to 430 Ma, the smaller continents drifted across the equator, while the northern hemisphere (NH) rain belt extended longitudinally and the southern hemisphere (SH) rain belt contracted. Then, the continents slowly reassembled and formed the single supercontinent Pangea around 250 Ma (
Fig. 1B). Beginning 170 Ma, Pangea broke up and drifted toward the North Pole. At around 80 Ma, the continental distribution was most fractured (
Fig. 1C), partly because of tectonic motions and sea level rise. Then, the continents slowly evolved to today’s configuration (
Fig. 1D), featured by the Atlantic expansion, the assembly of Eurasia, and the equatorward drift of Australia. The geographic distribution of tropical precipitation changed markedly with continental evolution. For example, the supercontinent Pangea was home to the intense rainfall zone on its eastern side, known as the Megamonsoon (
20,
26); the formation of the South Pacific Convergence Zone accompanied the continental drift of Australia. However, in all periods, the global distribution of precipitation featured a broad tropical rain belt with a peak in each hemisphere (fig. S2), which is the subject of this study: the ITCZ.
Here, we focus on migrations of the annual- and zonal-mean ITCZ while leaving other features of the ITCZ, such as its seasonality, for future study. We quantify the ITCZ position using three quantities (see Materials and Methods): the latitude of the tropical precipitation centroid (ϕ
ITCZ), defined as the area-weighted mean latitude of zonal-mean precipitation from 20°S to 20°N [e.g., (
27)]; the latitude of the tropical precipitation peak (ϕ
pp), defined as the latitude of the zonal-mean tropical precipitation maximum; and the precipitation asymmetry index (PAI), which quantifies the hemispheric asymmetry of tropical precipitation [e.g., (
28,
29)]. Time series of these three parameters are shown in
Fig. 2A, and the corresponding zonal-mean precipitation climatology are shown in fig. S3. The centroid, ϕ
ITCZ, shows relatively weak variability (ranging from 3°S to 4°N); the peak, ϕ
pp, shows large variability (9°S to 9°N) and sudden jumps between hemispheres, and the asymmetry index, PAI, varies from −0.3 (i.e., SH tropical precipitation is 30% stronger than NH tropical precipitation) to 0.36. Highlighting different aspects of the ITCZ, the three parameters are highly correlated. In the following, we mainly focus on ϕ
ITCZ for simplicity, keeping in mind that migrations of the ITCZ accompany systematic changes of atmospheric and oceanic circulations.
Aiming for a causal explanation of migrations of the ITCZ, we avoid linking the ITCZ position to internal climate variables, such as SST, because, then, one would need to answer the equally difficult question of what causes the SST changes in the simulations. Instead, we assess how ITCZ migrations are caused by changes in external parameters of the atmosphere-ocean climate system, such as concentrations of CO2, insolation, and continental evolution (see Materials and Methods for how these are specified in the simulations).
To achieve our goal, we adopt an energetic framework [e.g., (
3,
9)] to examine migrations of the ITCZ. Recognizing that the ITCZ is part of the rising branch of the time-mean tropical atmospheric meridional overturning circulation, this framework posits that ϕ
ITCZ collocates with the latitude of the energy flux equator (ϕ
EFE), which is the tropical latitude with zero vertically integrated meridional atmospheric energy transport. If one applies a linear approximation to the meridional distribution of atmospheric energy transport near the equator and assumes the slope to be nearly invariant with climate state, then −ϕ
EFE is proportional to the cross-equatorial atmospheric energy transport (
Fatm, positive values of which indicate northward transport). This energetic framework has been used to explain ITCZ variability from seasonal to millennial time scales [e.g., (
2,
3,
27)]. Our simulations confirm the close correlation among ϕ
ITCZ, ϕ
EFE, and
Fatm (
Fig. 2, A and B, and fig. S4). The correlation between ϕ
ITCZ and ϕ
EFE is 0.89, and that between ϕ
ITCZ and −
Fatm is 0.89. The sensitivity of ϕ
ITCZ to
Fatm is −3.3° PW
−1 (
Fig. 2D), which agrees well with that found over the seasonal cycle and in externally forced annual averages in observations and coupled climate models (
27). Furthermore, considering the energy balance of each hemisphere (fig. S5 and Materials and Methods), we may separate
Fatm and, thus, ϕ
ITCZ into components associated with cross-equatorial ocean heat transport and the hemispheric asymmetry of radiation
Here,
Focn is the ocean heat transport across the equator (positive denotes northward), and δ
R is the hemispheric asymmetry (NH minus SH) of net radiative energy input at the top of atmosphere (TOA).
Equation 1 states that either a northward cross-equatorial ocean heat transport (
Focn) or a positive net radiative heating asymmetry (δ
R) favors a NH ITCZ.
Now, we examine the time series of
Focn and δ
R (
Fig. 2C). From 540 Ma to present,
Focn shows a clear trend from negative to positive with the sign changing around 170 Ma, superimposed with variations on shorter time scales. Conversely, δ
R has a negative trend of similar strength; thus, their sum
Fatm shows no apparent linear trend over the whole time period. With the trend removed,
Focn and δ
R show no apparent correlation (the correlation coefficient of the detrended time series is −0.38).
Figure 2C also demonstrates that variations in
Fatm are not dominated by either component; rather,
Focn and δ
R are of equal importance and together produce the temporal variations of
Fatm (fig. S6). The strong cancelation between
Focn and δ
R is not a coincidence. In the following, we will examine the ocean heat transport and radiation asymmetry individually and argue that they are both caused by continental configuration changes over geological time scales.
Hemispheric asymmetry of radiation
The hemispheric asymmetry of radiation, δ
R, may be separated into a shortwave (δ
S, positive denotes inward energy) and longwave component (δ
L, positive denotes outgoing energy), respectively, so that δ
R = δ
S − δ
L (
Fig. 3, A and B). We find that δ
S dominates the variations in δ
R, with δ
L being much smaller. In addition, δ
L is anticorrelated with ϕ
ITCZ (
Fig. 3B), consistent with high-level clouds and water vapor in the ITCZ trapping longwave radiation. Thus, δ
S variations may serve as a first-order approximation of δ
R variations.
Because annual-mean insolation is nearly hemispherically symmetric (the difference is less than 1 W m
−2), δ
S has to arise from the interhemispheric difference of planetary albedo. We hypothesize that variations in land-ocean albedo contrast dominate variations in the interhemispheric difference of planetary albedo. The clear-sky net shortwave radiation at the TOA shows that land reflects more solar radiation than ocean [
Fig. 3 (C and D) shows two representative periods, and fig. S7 shows all periods]; clouds also affect planetary albedo, but we show later that they make little contribution to variations in the hemispheric asymmetry of radiation.
On the basis of these results, we construct a simple model of δS using only the landmass distribution. We simplify the planetary albedos over land and ocean as uniform (spatially and temporally over different geological periods) values of αp,lnd and αp,ocn, respectively. For each location, given its annual-mean insolation, we can calculate its local TOA net shortwave radiation depending on whether it is over land or ocean surface. Integrating it over each hemisphere, we have the net shortwave radiation of each hemisphere (see Materials and Methods) and, thus, its hemispherical asymmetry (δSes, the subscript denotes values estimated by the simple model).
We compare the calculation of the simple model to the simulation results to determine the coefficients of α
p,lnd and α
p,ocn. With each possible pair of α
p,lnd and α
p,ocn, we calculate the corresponding δ
Ses for each time period. The performance of the simple model may be estimated by the mean absolute difference between δ
Ses and the simulated δ
S by the Earth system model (see Materials and Methods). The discrepancy between δ
Ses and δ
S is minimized with a land-ocean albedo contrast of 0.1 (α
p,lnd − α
p,ocn = 0.1, corresponding to the white dashed line in
Fig. 3E), with little constraint on the absolute values of either albedo. This suggests that land-ocean albedo contrast is the main cause of δ
S. We may further constrain α
p,lnd and α
p,ocn by comparing the simple model–estimated and simulated global mean net shortwave radiation at the TOA (
Fig. 3F), yielding the optimal values of α
p,lnd = 0.39 and α
p,ocn = 0.29 (the asterisk in
Fig. 3F). These values are close to both observed estimates in modern climate [e.g., (
30)] and mean values diagnosed from all our simulations (the white triangle in
Fig. 3F).
Figure 3A shows that variations in δ
Ses estimated using these optimized albedos are very close to variations in δ
S and δ
R, with correlation coefficients of 0.90 and 0.96, respectively. Given the robustness of the parameters over geological time and the accuracy of the simple model, we speculate that it may be applied in other paleoclimate periods.
The dominant role of land-ocean albedo contrast in controlling δ
S is further demonstrated by additional lines of evidence. First, we note that the values of α
p,lnd and α
p,ocn obtained above represent planetary albedos (estimated at TOA), including effects of the atmosphere and surface. The atmosphere affects planetary albedo through absorption, scattering, and reflection of shortwave in clear and cloudy skies. Observational studies of modern climate indicate that the atmosphere contributes more to planetary albedo than the surface [e.g., (
31,
32)]. Thus, we expect the contrast between α
p,lnd and α
p,ocn to be smaller than the contrast between the surface albedos of land and ocean (typical values are 0.26 and 0.05, respectively) (
33), as is true of our estimated planetary albedos. Examining the effects of clear-sky and cloud processes on the hemispheric asymmetry of radiation, we find that the clear-sky component of the asymmetry dominates (fig. S8). The clear-sky planetary albedos over land and ocean diagnosed in our simulations (the white dots in
Fig. 3, E and F) are smaller than the all-sky planetary albedos [the white triangles in
Fig. 3 (E and F)], so using them in the simple model would overestimate global mean net shortwave radiation (
Fig. 3F). However, even those clear-sky values reproduce δ
Ses well (
Fig. 3E), highlighting the dominant role of land-ocean albedo contrast rather than the absolute values of albedos. Furthermore, there is strong cancelation between the longwave and shortwave effects of clouds on δ
R (fig. S8). Thus, while clouds play important roles in local and global-mean radiative balances, they seem to have little contribution to variations in the hemispheric asymmetry of radiation driven by continental drift.
Last, although the latitude of landmasses is part of the input parameters in the simple model, δ
Ses is mainly determined by the hemispheric difference in land area (δ
Alnd), as shown by the strong anticorrelation between δ
Ses and δ
Alnd (
Fig. 3A).
In summary, these results indicate that over geological time scales, land-ocean albedo contrast largely determines δR. Using only the horizontal distribution of land (and no information on surface type or orography), we can well estimate δR. This link between δR and continental configuration constitutes the first pathway for how continental evolution affects migrations of the ITCZ.
Ocean heat transport
Now, we examine the ocean heat transport, which, in our simulations, exerts an opposite influence on ITCZ migration compared with that of the hemispheric asymmetry of radiation. The cross-equatorial ocean heat transport plays an important role in setting the latitude of the ITCZ in the current climate (
3,
11,
34) and is dominated by the deep global meridional overturning circulation (GMOC) of the ocean (
35). This GMOC is clockwise (defined as northward at upper levels), mostly because of the Atlantic overturning circulation (
36). Cold and salty water sinks into the deep ocean in the North Atlantic, while the strong SH surface westerlies bring deep water to the ocean surface by Ekman pumping within the polar part of the Antarctic Circumpolar Current (ACC) (
37,
38). The clockwise GMOC transports heat from the SH to the NH and pins the ITCZ north of the equator. The even deeper overturning circulation associated with the deep-water formation around the Antarctica has a negligible contribution to heat transport (
39). Such bottom overturning circulation, if it exists, also has negligible contribution to ocean heat transport in all the simulated previous climates due to the small temperature difference between its upper and lower branches and is thus ignored and not counted in the GMOC in the context herein.
Consistent with the time series of
Focn (
Fig. 2C), our simulations show an anticlockwise GMOC (interhemispheric in almost all cases) before 170 Ma and a clockwise GMOC after 90 Ma, with oscillations in direction during the transition period (160 to 100 Ma; fig. S9). The dynamics of GMOC is a complicated, unsettled topic of great importance (
40,
41); it is out of our scope to provide a theory for the GMOC here. Instead, we boldly propose, on the basis of the simulation results, that for variability over geological time, the direction of the GMOC is mainly set by the hemispheric asymmetry of area-integrated wind stress on the ocean surface, especially in the middle latitudes. This assertion is supported by the strong anticorrelation (a coefficient of −0.86;
Fig. 4A) between
Focn and the hemispheric asymmetry of wind stress δ⟨τ⟩, defined as area-integrated wind stress over the ocean surface and normalized by hemispheric area in the NH minus that in SH (see Materials and Methods).
Prior studies based on seasonal to interannual variability of the modern ITCZ suggested that cross-equatorial ocean heat transport is dominated by the ocean’s shallow tropical circulation cells [e.g., (
42–
44)]. These studies argued that the trade winds exert stress on the tropical ocean, which, because of Sverdrup balance, couple the tropical shallow ocean cells with the tropical Hadley cell. However, decomposition of δ⟨τ⟩ (
Fig. 4A) shows that its variations due to tropical components (within 30°N/S) are close to zero, while its mid-latitude (30° to 70° in each hemisphere) component dominates. The component of δ⟨τ⟩ in the polar region is even smaller. This indicates that for ITCZ variability over geological time scales, the cross-equatorial ocean heat transport is determined by ocean deep overturning circulations driven by surface westerlies in middle latitudes rather than by tropical shallow cells driven by tropical winds.
Examining wind stresses (fig. S10) and the GMOC (fig. S9) in all the simulations confirms their strong relationship. Taking the 540-Ma period as an example (
Fig. 4, C and D), when landmasses are concentrated in the SH and the NH is a vast open ocean, there is a strong NH mid-latitude surface ocean current (
Fig. 4D), which may be analogous to the ACC in the present climate (although in the opposite hemisphere). This mid-latitude current is accompanied by deep-water upwelling (
Fig. 4C) due to Ekman transport and Ekman pumping induced by the wind stress and its curl and an anticlockwise GMOC. A similar relationship is found in other periods, such that the hemisphere with a wider ocean and stronger westerly winds in the mid-latitude is often the hemisphere with the rising branch of the GMOC (fig. S9 and S10). The wind-driven adiabatic component of the GMOCs dominates the diabatic component driven by vertical eddy diffusion (
40) in most periods, as inferred from the larger number of streamlines rising within the westerlies-driven upwelling zones than in other regions (fig. S9). This dominance of the adiabatic GMOC is consistent with the good relationship between
Focn and δ⟨τ⟩. This relationship is also consistent with zonal-mean theory that, for the present-day Earth, treats the net northward flux of ocean water by Ekman transport across the northern boundary of the ACC as a cause of the strength of the GMOC (the adiabatic part) (
45).
Because the mid-latitude component δ⟨τ⟩
mid dominates the total wind stress asymmetry δ⟨τ⟩, we further decompose δ⟨τ⟩
mid into components due to hemispheric differences in mid-latitude ocean surface area (δ
Aocn_mid) and due to hemispheric differences in mid-latitude wind stress intensity (see Materials and Methods). The component due to δ
Aocn_mid, which is a geographic parameter, explains most of the variability of δ⟨τ⟩
mid, while the component due to hemispheric differences in wind stress intensity is secondary (
Fig. 4B). This partition may be expected because the variability of δ
Aocn_mid due to continental drift over the examined time period is significantly larger than the variability of asymmetry of wind stress intensity. As a result, there is a very strong correlation (a coefficient of −0.91) between
Focn and δ
Aocn_mid. The greater importance of ocean area compared to wind stress intensity may allow for more faithful representation of climate in Earth system models because the GMOC in low-resolution models tends to be too sensitive to changes in wind stress (
46); the dependence of
Focn on δ⟨τ⟩ in our simulations is furthermore not due to a spurious sensitivity of the modeled GMOC to wind stress intensity. The results here do not argue against the roles of buoyance forcing in ocean dynamics, which is apparently important in setting the detailed structure of the GMOC (
47). However, among the simulations, the hemispheric difference of buoyancy forcing is arguably much smaller than the hemispheric difference of wind stress, and the buoyancy forcing is less deterministic in setting the direction of ocean circulation (
47). In summary, the above analysis indicates that
Focn is mainly set by the hemispheric asymmetry of mid-latitude ocean surface area due to the cross-equatorial ocean heat transport associated with the wind-driven GMOC.
Last, let us put the arguments made in the above two sections together. If we estimate δR by the simple model of radiation asymmetry and Focn by a linear fitting using δAocn_mid, the estimated Fatm follows the simulated Fatm reasonably well (fig. S11), especially for the longer time scale variability, albeit there are sizeable discrepancies in shorter time scales. Given that the only input for the above estimation is continental configurations (with coefficients retrieved by fitting simulations), this match further highlights the strong explanatory power of the simple arguments.