The Wayback Machine - https://web.archive.org/web/20210116133047/https://www.researchgate.net/profile/R_Bowyer/publication/272744459_Significance_of_Scent-Marking_by_Roosevelt_Elk/links/54eccaf60cf27fbfd771950e/Significance-of-Scent-Marking-by-Roosevelt-Elk.pdf